I think they're fitted every hundred metres or so, not just at the ends of the track circuit.
That makes sense to maintain occupancy throughout a section especially for leaf fall areas which can be many hundreds of metres long through cuttings.
The track circuit issue was "out there somewhere" with the 150s but didn't become really serious until the 158s came along - for a time they were only allowed out if paired with an older unit and some units were half a 158 with half a 156. The decision was then made to fit TCA to all Sprinters and Pacers but I think 158s were prioritised.
Shows how good the 158 and derivative designs were with regard to bogie, suspension and wheel profile design with consequent smooth ride characteristics. I've always liked them from my first ride.
It's noticable where they run to the exclusion of other designs that the wheel treads tend to track a very fine consistent path along the railhead, the rest of which, if not ground away by wheels of other trains remains heavily rusted. Therein lies a particular risk as if the contact point moves from its regular position for any reason, the wheels can easily be running on rust, which, if dry, can be a good insulator. An example of this is Exeter - Salisbury, normally only worked by 159s. Recent resignalling employed axle counters throughout however so there shouldn't be any problems in this reapect any more.
Incidentally automatic sanding systems were fitted most units from about 2000 onwards but were not allowed on four-axle units (153s and Pacers) until quite recently because of concern about dispensing too much sand and becoming invisible to track circuits.
Thanks for that info I was not aware of until now. Makes a lot of sense, but how is the risk controlled now? conductive sand?
As I said in another thread, track circuits are a nightmare and axle counters are a much better modern solution to train detection, but they still are susceptible to that awkward miscount problem when a wheel stops right above a sensor. It's always a right side failure (falsely occupied), but that's still a real problem, as operations are limited thence until the section can be reset safely. Nottingham seems a sensible compromise: Axle counters for the majority on plain line and through junctions; track circuits just through the long station platforms where split sections are provided to assist permissive movements and sharing.