First, current rail index applies only to DC motor EMU on third rail in SR zone.
The only such EMU left operational in that zone are 313.
456s and Southern 455s were the last previous types to go (SWT 455s retractioned with AC motors).
IGNORING things like Reps and 442s and 73s as bringing these in will only exponentially confuse matters :
In /very/ simple terms, the current rail index was count 1 for every EE507 motor in the train. Or, if you like, every coach in the train.
Hence a 4car = index 4 as /default/ types 4EPb 4Vep 4Cig etc had 4 motors - 319s were also nominally 4.
These operational values are at 675 (not 750) V DC at the third rail. Indeed all SR vaues quoted in the spotting books are 675 not 750 V.
In terms of kW, all such EMU operate with a current limit relay. If you want to know the maximum kW, power = volts x amps - it is 675 V x the current relay setting in Amp. I /think/ it is 400 A. So 675 V x 400 A = 270 kW max per motor.
Yes this is much greater than the nominal 250 hp quoted for an EE507 motor, but this is how DC EMU work, and perform so well, they can output much more power short term than the stated "trainspotter book" ratings.
In series only, it means 2 motors in series in SR EMU set up, so 135 kW per motor, much less than the nominal 250 hp. A 4car EMU would be maximum 4 x 135 = 540 kW in series only.
442 Reps 73s 71s 74s anything else not EE507 fitted have their own sums to do; each class unique in this respect.
None of this applies to AC motor EMU, which 377 375 450 etc etc are. These do not have conductor rail indices.
They work in very different ways, and, depending on who made it and when, there are differening software controls on current on the traction packs. Some of these (well for 700s at least) are activated by GSM by location. Not sure anyone could give a definitive statement.
The default 'country area' current is (I think) 4000 A, but, places like Seaford and Lymington, distance from substation makes things locally different; not sure the actual figures at these locations are in the public domain.
If I have got some values wrong, the principle is the same when someone else comes along and gives the right values.